Investigación y ciencia 346 - Julio 2005

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  • 7/23/2019 Investigacin y ciencia 346 - Julio 2005

    1/99

    GL@C_NL[@C [BURDM KB_BO_DM DJRAB_C[ FBJ_C[ KRDFWLKC[

    ARMLC ?992

    >-99 BR_C[

    _BWLFD[

    D_WLNLKLDMB[Y_CMLNB_DKLCF GB MD[ KMD@LGLD[1

    RF Y_COMB@D GB [DMRG YROMLKD

    KC@YRB[WC[ EDMCJBFDGC[

    BF MD FDWR_DMBQD

    KCML[LCFB[ GB D[WB_CLGB[

    BF BM [L[WB@D [CMD_

    KLKMC KBMRMD_

    BM _D\C

    < : : 9 ? 49 4 ; > 9 97

    9 9 ; 7 >

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    Gl`crffl{`c {budmkbrbordmMdrr| Kdelmm

    Bm kbrborc gbm ~drf glfflbrb gbm kbrborc gb md `abr bf crjdflqdklf | dkwl~lgdg&Md lf~b{wljdklf gb md{ ~drldklcfb{ co{br~dgd{ ycgrd mmb~dr d f wrdwd`lbfwcgb ydwcmcjd{ `bfwdmb{ b{ybkfflkc ydrd kdgd {buc&

    Amlc gb ?992

    F`brc ;7>

    ;EDKB&&&29- 499 | 429 dc{&

    7

    DYRFWB[Wbmbkc`flkdklcfb{&&&B~cmklf&&&Klbfkld fkmbdr&&&Bwlkd klbfwfflkd&&&N{lkd&

    ;?KLBFKLD\[CKLBGDGBkcmcjd ~bjbwdm&&&@lkrcb{wrldklf gbfwdrld | glbwd&&&Crgbfdgcrb{| dmjcrlw`c{ gb ~l{lf&&&KCF[WDF[- bm jbfgb md crdklf&

    ;5GBKB_KDBm mdjc Wdfjdflkd&

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    Djabrc{ fbjrc{ kfwlkc{Obrfdrg A& Kdrr | [wb~bf O& Jlgglfj{

    Tlq {b krbbf yrcfwc djabrc{ fbjrc{bf bm mdocrdwcrlc&

    4>

    _bwlfd{ drwlfflkldmb{Hzdobfd Ocdebf

    @bgldfwb `lkrcklrklwc{ kc`ydkwc{ tbrb`bgdf md b{wrkwrd fbrcfdm gbm kbrborcycgrdf ndorlkdr{b rbwlfd{ gb {lmlklc tbrb{wlw|df md ~l{lf- cac{ rcocwlqdgc{ | cwrc{{bf{crb{ lfwbmljbfwb{&

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    Klwctlfb{l{ bf kmmd{ bkdrlcwd{@brkbgb{ Ydrgc

    Bf md klwctlfb{l{- bwdyd fflfdm gbm klkmc kbmmdr- {b ncr`d fd odrrbrd tb {bydrdrbm `dwbrldm jbfwlkc rbklf gl~lglgc bf gc{ kmmd{ elad{&

    79

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    Bm rd|cAc{bye _& Gz|br

    Fb~d{ lf~b{wljdklcfb{ {ord|df md {cryrbfgbfwbkc`ymbalgdg gb md{ wcr`bfwd{ bmkwrlkd{& Bm rd|c

    {ljb ymdfwbdfgc yrcomb`d{ gb{kcfkbrwdfwb{&

  • 7/23/2019 Investigacin y ciencia 346 - Julio 2005

    3/99

    Edmcjbfrc{ crjflkc{Jcrgcf Z& Jrloomb

    Dftb {bmbf d{ckldr{b d md lfg{wrldt`lkd- mc{ kc`yb{wc{ edmcjbfdgc{

    dofgdf- wd`olf- bf fb{wrc bfwcrfc fdwrdm&

    52WDMMB_\MDOC_DWC_LCYfgmc jrd~l`wrlkc-ycr @drk Ocdgd

    55ARBJC[@DWB@WLKC[D`ljc{ ydrd {lb`yrb3-ycr Adf @&_& Ydrrcfgc

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    GL_BKWC_ JBFB_DM Ac{ @&d dmgbrd{ JdmmdrgcGL_BKWC_D NLFDFKLB_D Ylmdr Orcfkedm JdrnbmmdBGLKLCFB[ Adf Ybgrc Kd`yc{ J`bq

    Mdld Wcrrb{ Kd{d{Y_CGRKKLF @&d Krq Ljmb{ld{ Kdyf

    Dmobrw @drf Jdrd[BK_BWD_D Yrlfflkdklf @d|crdm @drwfbqDG@LFL[W_DKLF ^lkwcrld Dfgr{ Mdljmb{ld[R[K_LYKLCFB[ Kcfkbyklf Crbfb{ Gbmjdgc

    Cmjd Omdfkc _c`brcBGLWD Yrbf{d Klbfwfflkd- [&D& @fwdfbr- ;;< yrdm& 4&d959?4 Odrkbmcfd %B{ydd,Wbmncfc

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    LF^B[WLJDKLF\ KLBFKLD-amlc- ?992 ;

    &&&klfkbfwd dc{

    FRB^C BMB@BFWC&Grdfwb dmjfc{ gd{ d yrlfklylc{ gbb{wb dc eoc dmjc fb~c odac md mq gbm {cm- dftbfc bf gb`d{ldgd kdfwlgdg& T`lkc{ gb md Rfl~br{lgdggb Kdmlncrfld kcf{ljlbrcf krbdr 4: wc`c{ gbm bmb`bfwc494& Md {{wdfkld- odwlqdgd `bfgbmb~lc %{`ocmc @~8decrd @g, bf ecfcr gbm ydgrb gb md wdomd ybrlglkd- {bncr` oc`odrgbdfgc bm bmb`bfwc - {b gb{lfwbjrd ycr ffl{lf b{ycfwfbdkcf f {b`lybrcgc kc`yrbfglgc bfwrb wrblfwd `lfwc{| ~drld{ ecrd{&

    KD_OF& Bm dfml{l{ gbm kdrof d ydrwlr gb {{ kcf{wlw|bfwb{ bmb`bfwdmb{ fdgd fc{ glkb dkbrkd gb m kc`c

    {{wdfkld& Ydrd bmmc- fbkb{lwd`c{ kcfckbr { b{wrkwrdt`lkd- b{ gbklr- gb t `cgc {b bfmdqdf {{ wc`c{ydrd ncr`dr `cmkmd{& Bm b{nbrqc ~dmb md ybfd- yb{ kdfwc `{ {b {byd {corb k`c {bkcfncr`d- kcf `{ yrbkl{lfycgr gb{kc`ycfbr{b ydrdcowbfbr {{wdfkld{ t`lkd{gb lfwbr{& \d {b ed ybrnbkklcfdgc f yrckb{c tb ybr`lwbgb{kc`ycfbrmc bf `{ gb ?99kc`yb{wc{- dmjfc{ kc`ymbwd`bfwb fb~c{ c bf kdfwlgdgb{ffkd kcf{bjlgd{& B{ b{wb ebkec bm tb mmb~ d t`lkc{ |kcf{br~dklcfl{wd{ d kcfkmlrtb mc `{ gb{ylmndrrdgcr tbybgb edkbr{b kcf bm kdrofb{ tb`drmc&

    &&&klbf dc{

    BM KDFDM GB YDFD@&Bm yrb{lgbfwb _cc{b~bmw ed kc`ydrdgc mc{ gb{brwcrb{ tb edf~bmwc d kd{d gb{gb Ydfd`-kcf md ockd mmbfd gb krwlkd{ |

    tbad{- d mc{ yckc{ {cmgdgc{`bgrc{c{ | jrrmc{ tb {b bkedf dwr{ bf bm nrdjcrgb md odwdmmd& Fdglb {ycfb tb bm l{w`c gb Ydfd`{bd yrbkl{d`bfwb decrd f odmfbdrlc- f {dfdwcrlc flf mjdr gb ~dkdklcfb{8 dm yrcylc wlb`yc- { kc`lbfqdd ydrbkbr tb eol{b`c{ dkc`bwlgc mc{ wrdodac{ gbkcf{wrkklf {lf docrgdr mc{ yrbydrdwl~c{ b{ybkldmb{ ydrdmd rbkbyklf- dmcad`lbfwc | dml`bfwdklf gb md nbrqdmdocrdm- tb md{ `| ybfc{d{ kcfglklcfb{ bf bm l{w`cedkbf fbkb{drld{& [bd kc`c {bd- tb bm barklwc {bbfkdrjb gbm wrdodac& [brd wbfwdr dm gb{d{wrb bfkdrjdrgb mc{ klgdgc{ | md dml`bfwdklf d mc{ rb{ycf{domb{ gbmd{ kd{d{ gb e{ybgb{&

    GLD@DFWB[ D_WLNLKLDMB[& \d bf yc{b{lf gbm ecrfcbmkwrlkc- bm yrcnb{cr Ebfrl @cl{{df- b`lfbfwb t`lkcnrdfk{- lfwbfw rbyrcgklr bm yrckb{c tb {b {ycfb{ljb md fdwrdmbqd ydrd ncr`dr gld`dfwb{& Md cybrdklf b{bfkldm kcf{l{wb bf gl{cm~br bm kdrocfc bf elbrrc`dfwbflgc bf n{lf d ?:>9 cK& Bm {ljlbfwb yd{c- bfnrldror{kd`bfwb md `d{d ydrd tb ncr`b fd kc{wrd {mlgd&Mbjc- kdfgc bm lfwbrlcr {b bfnrd- wrdwd gb glmdwdr{b8ybrc b{w dyrl{lcfdgc ycr md kdyd buwbrfd& Ycr bnbkwc gbmd bfcr`b yrb{lf rb{mwdfwb- bm kdrocfc {b {bydrd gbmelbrrc bf ncr`d gb fflfc{ jrdfc{ gb gld`dfwb{ fbjrc{| wrdf{ydrbfwb{&

    EC@O_B \ KDODMMC& Fb{wrd ncwcjrdnd rbyrb{bfwd fjryc obmmd`bfwb kc`yb{wc ycr bm b{tbmbwc gb fkdodmmc | bm b{tbmbwc gb f ec`orb- rbklbfwb`bfwb

    buyb{wc{ bf bm @{bc D`brlkdfc gb El{wcrld Fdwrdm&Md l`djbf `b{wrd tb mc{ eb{c{ gb f ec`orb |

    f kdodmmc {cf rljrc{d`bfwbkc`ydrdomb{- {l olbf bm yrl`brckcf{br~d `{ gb md{ kdrdkwbr{wlkd{ yrl`lwl~d{ kc`fb{ dwcgc{ mc{ `d`nbrc{- `c{wrdfgc bm kdodmmc `d|cr b{ybkldmlqdklf bf md b{wrkwrd gb {{buwrb`lgdgb{ | { gbfwdgrdwrlwrdgcrd&

    &&&klbfwc klfkbfwd dc{

    DFKMD_ MD D_BFD&Yrckbgbfwb{ gb Ecmdfgd- {b edf rbklolgc bf md Cfflklfd gb Ydwbfwb{md{ {b`lmmd{ gb afkc `drlfc%Drfgc drbfdrld, | gb elbrodbrcybd gb md{ gfd{ %Bm|`{drbfdrl{,- tb edkb wlb`yc {bb`ymbdf bf dtbm yd{ ydrd gc`bdr mc{ k`mc{ gb drbfdbf md{ kc{wd{ `drlfd{& [b edfl`ycrwdgc b{d{ {b`lmmd{ ydrdbuybrl`bfwdrmd{ d mc mdrjc gbwcgd md kc{wd dwmfwlkd- gb{gb

    @dlfb ed{wd Nmcrlgd& Mc{ fwrlbfwb{ tb dycrwdf b{d{ elbrod{ fc od{wdf ydrd tb~dmjd md ybfd kmwl~drmd{- ybrc md kc`olfdklf gb d`od{rb{mwd gb jrdf wlmlgdg- yb{ wrdodf md{ drbfd{ {bmwd{ gbmd kc{wd `drlfd- dmqdfgc kcf bmmc fd odrrbrd ybrgrdombnrbfwb d md lf~d{lf gbm ckdfc {corb md{ kc{wd{& Gb ebkec-Ecmdfgd gbob { yrcyld bul{wbfkld- bf `bglgd kcf{lgbrdomb- d md dkklf yrb{br~dgcrd gb md{ `l{`d{&

    @DGB_D YD_D MD D_@DGD& Kfwc{ rocmb{ ncr`df fotb3 [b fbkb{lwdf ?999 rocmb{ dgmwc{- c md kc{bked`dgrd gb 45 ebkwrbd{ gb oc{tb- ydrd {`lfl{wrdr bm`dgbrdab gb f {cmc otb gb :7 kdcfb{&

    B{tbmbwc{

    gb ec`orb

    | kdodmmc

    kc`ydrdgc{-

    4

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    7 LF^B[WLJDKLF\ KLBFKLD-amlc- ?992

    DYRFWB[

    WBMBKC@RFLKDKLCFB[

    Nlord bf kd{dFc b{w {lbfgc nklm korlr wd`olf kcf fflord ywlkd- bf ~bq gb kcf kcorb- b{d mwl`d `lmmdtb {bydrd mc{ ecjdrb{ gb md{ mfbd{ ywlkd{ tb ncr`df md{ rbgb{ gb wbmbkc`flkdklcfb{&Md kdydklgdg gb wrdf{`l{lf gb md fflord b{ `ke{l`c `d|cr tb md gbm kcorb1 bf bm mdocrdwcrlc |d kdfdmlqd glbq wbrdolw{ %glbq olmmcfb{ gb olw, ycr {bjfgc- | ycgrd mmbjdr ed{wd mc{ klbfwbrdolw{- `lbfwrd{ tb fd kcfbulf gb DG[M tb wlmlqd bm elmc gb kcorb wbmbnflkc {mc dmkdfqd fc{ `lmmcfb{ gb olw{ ycr {bjfgc& Bm lfkcf~bflbfwb ed{wd decrd edod {lgc bm kc{wb& [lfb`odrjc- bm yrbklc gb kdombdr kcf fflord fd kd{d bf md{ fb~d{ rodflqdklcfb{ ed gl{`lflgced{wd ljdmdr{b dm gb md lf{wdmdklf gbm elmc gb kcorb& ^brlqcf Kc``flkdwlcf{ ymdfbd cnrbkbrdkkb{c d Lfwbrfbw ycr fflord bf B{wdgc{ Rflgc{ dm `bfc{ d wrb{ `lmmcfb{ gb ecjdrb{ ydrd fflfdmb{ gb ?992& Gl~lgb md {bdm gb fd mfbd gb fflord gb >?9 `bjdolw{ ycr {bjfgc bf ;? mfbd{gb docfdgc{- kdgd fd gb 4

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    LF^B[WLJDKLF\ KLBFKLD-amlc- ?992 2

    BWLKD KLBFWLNLKD

    Lf{c{ybkedgd `djflwg gb md{ lrrbjmdrlgdgb{ klbfwfflkd{

    Bf ?999- f crjdfl{`c gbm jcolbrfc b{wdgcflgbf{b kcf{lgbrdod dkwc{ gb `dmd kcfgkwd klbfwfflkd d`ddr- lf~bfwdr c ymdjldr& Md{ lf~b{wljdklcfb{- od{wdfwb ml`lwdgd{- tb {bedf rbdmlqdgc kcf md lfwbfklf gb d~brljdr bm dmkdfkb gb b{wc{nrdgb{ drrcadf fd lfklgbfkld gbm fc c gc{ ycr klbfwc& Bf?99?- lf{wdfkld{ gb md dg`lfl{wrdklf yrcy{lbrcf- wd`olf bfB{wdgc{ Rflgc{- f b{wglc `{ d`ymlc tb w~lb{b bf kbfwdcwrc{ wlyc{ gb yrkwlkd{ rbyrcodomb{- ybrc md{ lf{wlwklcfb{ kcmbjldmb{ gb md{ rbd{ olcmjlkd{ | `glkd{ {b fbjdrcf d mmb~drmcd kdoc& Decrd- bf aflc- wrb{ lf~b{wljdgcrb{ edf yomlkdgc bfFdwrb mc{ rb{mwdgc{ gb fd{ bfkb{wd{ dffl`d{ rb`lwlgd{d ;>99 klbfwfflkc{ tb rbklolbrcf ycr yrl`brd ~bq ncfgc{ ydrdf yrc|bkwc gb lf~b{wljdklf bfwrb 4 ycr klbfwc kcfnb{ tbfc edod yrb{bfwdgc gdwc{ tb kcfwrdgbkdf {{ lf~b{wljdklcfb{dfwbrlcrb{- bm 42-2 tb edod kd`oldgc bm gl{bc c mc{ rb{mwdgc{ gb f buybrl`bfwc ycr yrb{lcfb{ gb f ydwrcklfdgcr | bm4?-2 tb edod ebkec md ~l{wd jcrgd dm {c ycr cwrc{ gb gdwc{gbnbkwc{c{& Bf wcwdm- md wbrkbrd ydrwb ;5 ycr klbfwc gbm yrl`brjryc- ?2 gbm {bjfgc dg`lwd edobr kc`bwlgc dm `bfc{ fdgb glbq lrrbjmdrlgdgb{ tb- gb {br gb{kolbrwd{- {bjrd`bfwb mb{ycfgrdf fd {dfklf dm rb{ycf{domb8 ybrc dgb`{ f 4;-2 ycrklbfwc glkb edobr rbkrrlgc d gl{bc{ buybrl`bfwdmb{ gbfflklbfwb{| f 42-; tb ed {yrl`lgc gdwc{ ycr fd {bf{dklf ~l{kbrdm&Mc{ dwcrb{ gbm wrdodac `dflfflb{wdf kfwc mb{ edf l`yrb{lcfdgcmc{ rb{mwdgc{ gb md bfkb{wd& Bm glrbkwcr gbm Kbfwrc gb Olcwlkdgb md Rfl~br{lgdg gb Ybff{|m~dfld- bf kd`olc- gbkmdrd d Fdwrbtb fc gbobf edkbr ybf{dr tb md klbfkld b{wd kcrrcgd& [lf

    b`odrjc- km b{ md rco{wbq- kc`c glkbf mc{ b{wdg{wlkc{- gbmd ~dmlgbq gb f rb{mwdgc klbfwfflkc dfwb md{ kcfgkwd{ ggc{d{tb ybgdf lfwbr~bflr bf { cowbfklf3 Kfwc{ rb{mwdgc{yomlkdgc{ | dg`lwlgc{ fc {cf gb ~brgdg ffldomb{3

    KLBFKLD FRKMBD_

    Rfd n{lf rbdm`bfwb nrd

    Fcr`dm`bfwb- md n{lf fkmbdr rbtlbrb f kdmcr gb `kec{ `lmmcfb{ gb jrdgc{&Bul{wb- {lf b`odrjc- f gl{yc{lwl~c gbm wd`dc gb f odmgb tb nfgb fkmbc{ d wb`ybrdwrd{ lfnbrlcrb{ d md gb kcfjbmdklf gbm djd& Bm bmb`bfwc kbfwrdmgb md `tlfd bf kb{wlf b{ f krl{wdm gb wdfwdmdwc gb mlwlc- tb jbfbrd f kd`ycbmkwrlkc kdfgc {b kdmlbfwd& Rfd djad gb wfj{wbfc- tb {b kcfbkwd d md {{wdfkldylrcbmkwrlkd bfnrldgd ycr `bglc gb flwrjbfc mtlgc- bfnckd b{b kd`yc gb wdm `cgc

    tb kdfgc bm krl{wdm {b kdmlbfwd ed{wd fc{

    ; jrdgc{- mc{ lcfb{ gb gbwbrlc tbrcgbdf d md djad {dmbf l`ym{dgc{ d ~bmcklgdg {fflklbfwb ydrd tb {b n{lcfbf bff omdfkc kdrjdgc wd`olf kcf gbwbrlc& Mc{ jbfbrdgcrb{ gb bfbrjd ycr n{lfylrcbmkwrlkd {cf fd tl`brd1 f {cmc klkmc gb md `tlfd yrcgkb dmrbgbgcr gb49 fdfcacmb- bfbrjd tb dybfd{ od{wd ydrd kdmbfwdr fd{ yckd{ `lmmcf{l`d{ gbjrdgc bm djd kcfwbflgd bf f gbgdm& Df d{- bm yrckb{c ycgrd mmb~drfc{ ed{wd jbfbrdgcrb{ gb fbwrcfb{ gb wd`dc gb ocm{lmmc- `kec `{ {l`ymb{ tb mc{ dkwdmb{-tb ~dmgrdf kc`c yrcym{crb{ bf d{wrcfd~b{ gb `lfldwrd c ydrd wbrdyld{ `glkd{gb ~dfjdrgld&

    Kedrmb{ T& Kecl

    Mq gb n{lf& Rf omdfkc gb ym{wlkc gb ?-2

    kbfw`bwrc{ gb dfkec kdrjdgc kcf gbwbrlc

    orlmmd bf md c{krlgdg kdfgc {corb m lfklgbf

    lcfb{ gb gbwbrlc&

    NL[LKD

    Kcfjbmdklf jrdfmdr

    Kdfgc md `dwbrld- bf md `d|crd gb {{ ncr`d{- {b kdmlbfwd- {b ~bm~b`{ lgd& Ybrc md{ `d{d{ jrdfmc{d{ ydrbkbf fd bukbyklf& OcoOberlfjbr | Hdrbf Gdflbm{- gb md Rfl~br{lgdg Ghb- mmbfdrcf gb kdflkd{gb ym{wlkc fd wcm~d klmfgrlkd | kbrrdgd& Bf rbyc{c- ydrbkdf f {mlgckrl{wdmlfc- kc`c bm elbmc& Kdfgc {b md{ klqdmmdod jlrdfgc md ydrwb {ybrlcrgbm rbklylbfwb- md bfbrjd tb {b mb{ kbgd d{ nfgd bm krl{wdm ed{wdf b{wdgc `{ d`crnc | lgc- kc`c brd gb b{ybrdr& Ybrc dm lf|bkwdr`{ bfbrjd `bgldfwb ~lordklcfb{ gb md wcm~d- ckrrd dmjc buwrdc1 {brbkcfjbmdodf& Rf jcmybwbc lfwbf{c wcrfd md{ kdflkd{ `c`bfwfbd`bfwb lfjr~lgd{- mc tb mb{ ybr`lwb dyrbwdr{b `{ fd{ d cwrd{- {bjf mdelywb{l{ kcf tb d`oc{ lf~b{wljdgcrb{ tlbrbf buymlkdr bm nbf`bfc& Bmrb{mwdgc kcfwrdglkb md rdqf ycr md tb {b {df dml`bfwdgcrb{ ~lordwcrlc{bf bm yrckb{d`lbfwc gb ycm~c{ | jrdfmdgc{- d {dobr- tb md djlwdklfnd~crbkb md lgbq gb mc{ jrdfc{&

    A& _& @lfhbm

    Md{ kdflkd{ {c`bwlgd{ d ~lordklf `b{wrdf f kc`ycrwd`lbfwc d yrl`brd ~l{wd ydrdgalkc& O_LDF

    FD_DFAC

    RKMD

    %drrlod,8KC_WB[LD

    GB

    OCOO

    BE_LFJB_

    Rfl~br{lgdggbGhb%dodac,

  • 7/23/2019 Investigacin y ciencia 346 - Julio 2005

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    > LF^B[WLJDKLF\ KLBFKLD-amlc- ?992

    Bf f gd jrl{- d `bgldgc{ gb bfbrc- Mdzrbfkb[``br{- rbkwcr gb md Rfl~br{lgdg gb Edr~drg-

    gba kdbr tb bfwrb md{ kd{d{ gb md b{kd{bq gb`abrb{ gbglkdgd{ d md{ klbfkld{ ycgrdf kcfwdr{b

    fd{ glnbrbfkld{ lffdwd{ bf md kcf{wlwklf gb mc{ kbrborc{`d{kmlfc | nb`bflfc& B{d{ gbkmdrdklcfb{ dwlqdrcf bmnbjc gb f gbodwb tb mmb~d e`bdfgc gb{gb edkb f{ljmc- gb{gb tb dmjfc{ `bglgcrb{ gbm kbrborc e`dfc{b ~dmlbrcf gb fc gb mc{ rb{mwdgc{ tb edodf cowbflgc-tb bm wd`dc `bglc gbm kbrborc gb md{ `abrb{ b{ `bfcr tb bm gb mc{ ec`orb{- ydrd rdwlnlkdr md {ybrlcrlgdglfwbmbkwdm gb {wc{&

    Fdglb- ed{wd md nbked- ed gb{kolbrwc yrbod dmjfdgb tb md `abr- ycr {{ ybkmldrlgdgb{ dfdw`lkd{- {bdlfkdydq gb dmkdfqdr md bukbmbfkld bf `dwb`wlkd{- n{lkd

    GL@C_NL[@C [BURDM

    [ML@

    NLM@[

    Bm kbrborc gbm ~drf

    glfflbrb gbm kbrborc gb md `abrbf crjdflqdklf | dkwl~lgdg&

    Md lf~b{wljdklf gb md{

    ~drldklcfb{ co{br~dgd{ ycgrd

    mmb~dr d f wrdwd`lbfwc

    gb ydwcmcjd{ `bfwdmb{

    b{ybkfflkc ydrd kdgd {buc

  • 7/23/2019 Investigacin y ciencia 346 - Julio 2005

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    LF^B[WLJDKLF\ KLBFKLD-amlc- ?992 :

    c lfjbflbrd %~d{b bm rbkdgrc Md ycm`lkd gbm rbkwcr gb Edr~drg,& [b ed gb`c{wrdgc tb bm kbrborc gb

    f {buc {b ydrbkb `kec bf `mwlwg gb d{ybkwc{ dmgbm cwrc& Fc co{wdfwb- d mc mdrjc gb mc{ glbq mwl`c{dc{- md{ lf~b{wljdklcfb{ rbdmlqdgd{ {corb fd ~drlbgdg gb kb{wlcfb{- tb ~df gb{gb bm yrckb{d`lbfwc gbmmbfjdab- md kdydklgdg gb crlbfwdr{b bf f rbkcrrlgcc bm rbjl{wrc gb rbkbrgc{ b`cwl~c{- edf {dkdgc d mdmq f d{c`orc{c dodflkc gb ~drldfwb{ b{wrkwrdmb{-t`lkd{ | nfklcfdmb{ bfwrb bm kbrborc gbm ~drf |bm gb md `abr&

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    KB_BO_DM MD__\ KDELMM

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    5 LF^B[WLJDKLF\ KLBFKLD-amlc- ?992

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    LF^B[WLJDKLF\ KLBFKLD-amlc- ?992 -YDJ[&7:7:?44>9?8 ?994& ebyye#949>?2- wrdodac F&c 4?79?:8 ?99?& ebyye#944??7:

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  • 7/23/2019 Investigacin y ciencia 346 - Julio 2005

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  • 7/23/2019 Investigacin y ciencia 346 - Julio 2005

    29/99

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    30/99

    ?5 LF^B[WLJDKLF\ KLBFKLD-amlc- ?992

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  • 7/23/2019 Investigacin y ciencia 346 - Julio 2005

    38/99

    ;> LF^B[WLJDKLF\ KLBFKLD-amlc- ?992

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  • 7/23/2019 Investigacin y ciencia 346 - Julio 2005

    39/99

    LF^B[WLJDKLF\ KLBFKLD-amlc- ?992 ;:

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  • 7/23/2019 Investigacin y ciencia 346 - Julio 2005

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  • 7/23/2019 Investigacin y ciencia 346 - Julio 2005

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    LF^B[WLJDKLF\ KLBFKLD-amlc- ?992 22

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